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Abstract Glutathione (GSH) is the main non-protein thiol in cells whose functions are dependent on the redox-active thiol of its cysteine moiety that serves as a cofactor for a number of antioxidant and detoxifying enzymes

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The presence of ascorbate in the apoplast is well characterised, along with an apoplastic enzyme ascorbate oxidase, which affects its oxidation state [32]

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