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The WWC also plays a role in regulating the oxidation state of the chloroplast-targeted 2-Cys PRXs, which, together with specific atypical TRXs such as ACHT1ACHT4 and TRXL2, are involved in the transfer of oxidative equivalents from H 2 O 2 to target chloroplast proteins, such as those of the reductive pentose pathway (Ojeda et al